The crow king is dead, and in the field below all the creatures tremble as the Murder gathers to choose a new king from the rival sons Sintus, Milus and Nascus. When the crows drive everyone from the field to keep the reckoning secret, the quail Ysil, Cormo and Harlequin believe they must simply follow their elders to safety.

But when the crows turn against each other, the forest becomes full of danger. In the confusion the last wolf, Asmod, shucks off his isolation and begins to raise an army to claim the kingdom for his own. As hidden truths are brought to light and enormous sacrifices are made Ysil and his friends must make an epic journey and an unthinkable alliance if the lesser animals are to survive.